Pollination: Male and female gametes in flowering plants develop in the pollen grain and embryo sac. Because both types of gametes are immotile, they must be brought together before fertilisation can occur, which is accomplished through a process known as pollination. Pollination is the transfer of pollen grains shed from the anther to the stigma of the pistil.
Flowering plants have evolved many adaptations to ensure effective pollination. Pollination is classified into three types based on the pollen source. This article delves into the important topic of pollination in Class 12, specifically sexual reproduction in flowering plants , including pollen-pistil interactions and techniques such as artificial hybridisation.
Pollination is the transfer of pollen from a plant's male reproductive organs to its female reproductive organs, which are usually the stigma of a flower. Pollen, which contains the genetic information required for plant reproduction, can be transferred within the same flower (self-pollination) or between flowers of the same species (cross pollination). This process is necessary for the production of seeds, which are required for the survival of plant species. Pollen can be spread through a variety of means, including water, wind, and animal pollinators like bats, insects, and birds.
Pollination is necessary for all flowering plants to reproduce. Pollination is classified into two types: self-pollination and cross-pollination. Self-pollination occurs when a plant has both male and female reproductive organs and pollinates its own flowers or plants. Cross-pollination occurs when an external agent transfers pollen from the male to the female reproductive organs, either between different plants or between flowers of the same plant. Cross-pollination increases genetic diversity because the offspring inherit traits from two different parent plants in the same species.
Pre Fertilisation - Structures And Events
When pollen moves from the anthers of a flower to the stigma of the same flower, it's called self-pollination. This pollination is common in plants with both male and female parts in the same flower.
Self-pollinating plants rely less on external factors for pollination, often using wind or small insects. The anthers and stigma are typically similar in length to aid pollen transfer in these flowers. Self-pollination can be of two types:
Sexual Reproduction in Flowering Plants MCQ
Cross pollination occurs when pollen is transferred from one flower's anther to another flower's stigma, usually between genetically different flowers. Birds, animals, water, wind, or insects are all examples of external pollinators. Depending on the agent, cross pollination can be classified as follows.
Hydrophilous Flowers: Pollinated by water, with small, inconspicuous flowers designed to hold pollen in place.
Zoophilous Flowers: Pollinated by animals, with pollen designed to stick to animal bodies for transfer.
Anemophilous Flowers: Pollinated by wind, with light, non-sticky pollen grains easily carried by the wind.
Entomophilic Flowers: Pollinated by insects, often with attractive features like bright petals and fragrance to attract insects. Pollen grains may be spiny or have extensions to stick to insects' bodies.
Ornithophilous Flowers: Pollinated by birds, a rare form of pollination.
Pollination is an important step in the reproduction of flowering plants. The transfer of pollen grains from the anther (male part) to the stigma (female part) of the same or another flower results in fertilisation. Plants use a variety of pollination agents, both biotic (living) and abiotic (non-living).
Wind pollination is more common in plants that use abiotic agents for pollination.
Pollination by water is rare in flowering plants and is limited to certain genera, mostly monocots. Plants like Vallisneria, Hydrilla, and marine sea-grasses use water for pollination. Water is a regular means of transporting male gametes in lower plants like bryophytes and pteridophytes.
The following are the plants that use water as an agent for pollination:
Animals serve as pollinators for the vast majority of flowering plants. Common animal pollinators include bees, butterflies, flies, beetles, birds, and bats. Insects, particularly bees, are the primary biotic pollinators.
Pollination is a complex process involving various agents and adaptations in plants. Understanding pollination mechanisms is crucial for the conservation and propagation of plant species.
Hermaphrodite flowers can undergo self-pollination, a process where pollen from the same flower fertilizes its ovules. However, continued self-pollination can lead to a phenomenon known as inbreeding depression. Inbreeding depression reduces the biological fitness of offspring produced by closely related individuals, leading to lower survival rates and fertility.
To avoid the negative effects of self-pollination, flowering plants have evolved various mechanisms to encourage cross-pollination, where pollen is transferred between different flowers. These mechanisms, known as outbreeding devices, include:
Monoecious: These plants have both male and female flowers on the same plant, such as castor and maize. While this arrangement prevents self-pollination (autogamy), it does not prevent pollination between flowers of the same plant (geitonogamy).
Dioecious: In some species, male and female flowers are present on separate plants, meaning each plant is either male or female (dioecy). This condition prevents both autogamy and geitonogamy.
In many bisexual flowers, the stamens (male reproductive organs) and carpels (female reproductive organs) do not mature at the same time. This temporal separation ensures that the pollen is released before the stigma (part of the carpel that receives pollen) becomes receptive, or vice versa, preventing autogamy.
In certain species, the anthers (part of the stamen that produces pollen) and stigma are positioned at different heights within the flower, preventing self-pollination by ensuring that pollen cannot reach the stigma of the same flower.
This genetic mechanism prevents self-pollen (either from the same flower or other flowers of the same plant) from fertilizing the ovules. It does so by inhibiting pollen germination or pollen tube growth in the pistil (female reproductive organ).
Pollen-pistil interaction is a crucial process in plant reproduction, encompassing all events from the deposition of pollen on the stigma (pollen-receptive part of the pistil) to the entry of the pollen tube into the ovule (structure containing the female gametophyte). This dynamic interaction involves the recognition of compatible pollen by the pistil, followed by either acceptance or rejection of the pollen.
The ability of the pistil to recognize pollen as compatible or incompatible is due to a continuous dialogue between the chemical components of the pollen and those of the pistil. If the pollen is compatible, the pistil promotes post-pollination events that lead to fertilisation. In contrast, if the pollen is incompatible, the pistil rejects it by preventing pollen germination or pollen tube growth.
After compatible pollination, the pollen grain germinates on the stigma, producing a pollen tube that grows through the tissues of the stigma and style (elongated part of the pistil) and reaches the ovary. In some plants, male gametes are formed during the growth of the pollen tube in the stigma, while in others, the pollen tube carries the male gametes from the beginning.
Once the pollen tube reaches the ovule, it enters through the micropyle (small opening in the ovule). Then it penetrates one of the synergids (cells in the ovule) through the filiform apparatus. The tip of the pollen tube bursts, releasing its contents, including the male gametes, inside the synergids. This leads to fertilisation and the development of the male gametophyte, ultimately resulting in seed formation.
Overall, pollen-pistil interaction is a dynamic process that can be manipulated by plant breeders to achieve desired hybrids, even in cases of incompatible pollinations.
Artificial hybridisation is a method used in crop improvement programs to control pollination and achieve desired traits in offspring. It involves several steps:
Double fertilisation is a unique process that occurs in angiosperms, where two fertilisation events take place during the same reproductive cycle, leading to the formation of two different structures:
Syngamy + Triple Fusion = Double Fertilisation
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